personal i would have the WWW/DNS/Email on one box
and make sure the ports on your router are being forward to the right PC IP address
and the firewalll is have them ports open for the services that are on each box
than try a site like canyouseeme.org
and do an port test to see if ur port 80 / 25 / 110 / 53 (u said ur DNS is working outside)
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